This was the first "Electric Eye" Plates (as listed by Johl) Electric Eye Plates
Electric Eye - In 1933, starting with the two-cent stamp, the Bureau of Engraving and Printing began experiments to improve the perforation process. The plate block at right is from one of the first electric eye plates ever printed. Note the position of the plate number, it's not in the corner as rotary press plate numbers normally are. The electric eye makes use of a photovoltaic cell employing a beam of light to aid in perforation accuracy. Although the experimental sheets had a series of dashes down the very center of the sheet and a single dash in the center of the left margin, the electric eye markings we are familiar with were not added to the sheet margins until 1939. The philatelist may note that many stamps after 1940 always seem to be well-centered. The electric-eye had a large part in this.
Electric Eye Reference ->
http://www.1847usa.com/Glossary/E.htm