No idea - they do not indicate something was splashed on them as they are too perfectly circular. They are all slightly different from each other. The colour does appear to be similar to the colour in the horse's collar when the images are enlarged.
The dots are not part of the original design, they were applied somewhere in the mail stream by USPS. I don't recall ever seeing any articles about the purpose, but it was before the USPS started using sprayed cancels. Testing? Coincidence?
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